Handsome Jack sends in waves of security forces in order to protect his daughter, but ultimately, Roland and the Vault Hunters, with the help of late arrival Lilith, successfully kill Angel. She requests to be killed in order to stop her father from charging the Vault Key and gaining control of The Warrior. Roland and the Vault Hunters assault the Hyperion compound and meet Angel, who is not only a real person, but a siren, and Handsome Jack's daughter. During the process, Mordecai's pet bird Bloodwing is captured and killed by Handsome Jack an enraged Mordecai vows revenge and joins the other three former Vault Hunters at Sanctuary.
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The Vault Hunters embark upon a series of missions in order to assemble what is needed to overcome these defenses, including obtaining an upgrade for Claptrap, visiting the Hyperion city of Opportunity, and enlisting the aid of Brick, now leader of the Slab clan of bandits. She also shares information about the formidable defenses of the facility, but insists that Lilith be kept as far away from the facility as possible. Angel communicates with the group in Sanctuary, and despite hostile treatment from Roland, divulges that the Vault Key is kept with her in a Hyperion facility. Jack's plan is to open Pandora's second Vault and unleash The Warrior, a powerful lava, eridium, and rock creature controlled by whoever releases it knowing that the key only charges itself once every 200 years, he uses Angel to forcefully charge the Vault Key. Meanwhile, rumors of an even larger Vault hidden on Pandora spread across the galaxy, drawing a new group of Vault Hunters to the planet in search of it. Now, Handsome Jack rules over the inhabitants of Pandora with an iron fist from his massive satellite built in the shape of an 'H', always visible in the sky in front of Pandora's moon. Handsome Jack, the president of the Hyperion Corporation, secures this new resource and makes use of it to attempt to 'bring peace' to the planet. After defeating The Destroyer, a valuable mineral called 'Eridium' started flourishing through Pandora's crust.
Instead, these Vault Hunters were confronted by an alien abomination known as 'The Destroyer' inside the Vault.
Five years have passed since the events of Borderlands, when the four Vault Hunters, Roland, Mordecai, Lilith, and Brick were guided by a mysterious entity known as 'The Guardian Angel' on the planet of Pandora to the Vault, a mysterious alien structure that was rumored to hold ancient and exotic technology and riches.
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